Arthur Middleton

Associate research scientist

Publications (BETA)

This is a PARTIAL listing of recent publications, and it will continue to grow as we populate our publications database.


  • Journal Article
    Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, D.E. McWhirter, and K. Gerow. “Native prey distribution and migration mediates wolf (Canis lupus) predation on domestic livestock in the Greater Yellowstone Ecosystem.” Canadian Journal of Zoology 94.4 (2016): 291-299. DOI: 10.1139/cjz-2015-0094
  • 2013

  • Journal Article
    Middleton, A.D.; Kauffman, M.J.; McWhirter, D.E.; Jimenez, M.D.; Cook, R.C.; Cook, J.G.; Albeke, S.E.; Sawyer, H.; White, P.J. “Linking anti-predator behaviour to prey demography reveals limited risk effects of an actively hunting large carnivore.” Ecology Letter 16.8 (2013): 1023-1030. DOI: 10.1111/ele.12133